A penguin's feet are nearly as cold as ice, while the bird's core body temperature is much higher. Which of the following explains this observation? 1) Penguins have a unique circulatory system that keeps their feet cold. 2) Penguins have a thick layer of blubber on their feet to insulate them from the cold. 3) Penguins have a high metabolic rate that generates heat in their core body. 4) Penguins have a specialized heat exchange system in their feet.